Well, I misremembered half of it, but Homestar Runner was one of the many flash animations of the flash-craze heyday. Occasionally, they were puppets. From nearly 25 years ago: https://m.youtube.com/watch?v=tLSgRzCAtXA

It’s probably turtles all the way down, but I think “A[n] British animators’s email segment co-star mispronounces misspells 90s greeting” might be an even smaller domino.

Edited to replace failing memory.

And corncream-pilled